Natural selection acts solely by accumulating slight successive favorable variations, it can produce no great or sudden modification; it can act only by very short steps.

When the sexes differ in beauty, in the power of singing, or in producing what I have called instrumental music, it is almost invariably the male which excels the female.

To my deep mortification my father once said to me, "You care for nothing but shooting, dogs, and rat-catching, and you will be a disgrace to yourself and all your family."

As the sense of smell is so intimately connected with that of taste, it is not surprising that an excessively bad odour should excite wretching or vomitting in some persons.

The fact of evolution is the backbone of biology, and biology is thus in the peculiar position of being a science founded on an improved theory, is it then a science or faith?

A novel according to my taste, does not come into the moderately good class unless it contains some person whom one can thoroughly love - and if a pretty woman, all the better.

We can allow satellites, planets, suns, universe, nay whole systems of universes, to be governed by laws, but the smallest insect, we wish to be created at once by special act.

The impossibility of conceiving that this grand and wondrous universe, with our conscious selves, arose through chance, seems to me the chief argument for 
the existence of God.

It is so easy to hide our ignorance under such expressions as the plan of creation or unity of design, etc., and to think that we give an explanation when we only restate a fact.

At some future period, not very distant as measured by centuries, the civilized races of man will almost certainly exterminate, and replace the savage races throughout the world.

...I have always maintained that, excepting fools, men did not differ much in intellect, only in zeal and hard work; and I still think there is an eminently important difference.

It strikes me that all our knowledge about the structure of our Earth is very much like what an old hen would know of the hundred-acre field in a corner of which she is scratching.

...I feel most deeply that the whole subject is too profound for the human intellect. A dog might as well speculate on the mind of Newton.— Let each man hope & believe what he can.—

Nothing is easier than to admit in words the truth of the universal struggle for life, or more difficult--at least I have found it so--than constantly to bear this conclusion in mind.

Formerly Milton's Paradise Lost had been my chief favourite, and in my excursions during the voyage of the Beagle, when I could take only a single small volume, I always chose Milton.

With highly civilised nations continued progress depends in a subordinate degree on natural selection; for such nations do not supplant and exterminate one another as do savage tribes.

On seeing the marsupials in Australia for the first time and comparing them to placental mammals: “An unbeliever . . . might exclaim 'Surely two distinct Creators must have been at work'”

Our ancestor was an animal which breathed water, had a swim-bladder, a great swimming tail, an imperfect skull & undoubtedly was an hermaphrodite! Here is a pleasant genealogy for mankind.

I have deeply regretted that I did not proceed far enough at least to understand something of the great leading principles of mathematics, for men thus endowed seem to have an extra sense.

About weak points [of the Origin] I agree. The eye to this day gives me a cold shudder, but when I think of the fine known gradations, my reason tells me I ought to conquer the cold shudder.

I suppose you are two fathoms deep in mathematics, and if you are, then God help you. For so am I, only with this difference: I stick fast in the mud at the bottom, and there I shall remain.

Man himself cannot express love and humility by external signs, so plainly as does a dog, when with drooping ears, hanging lips, flexuous body, and wagging tail, he meets his beloved master.

A grain in the balance will determine which individual shall live and which shall die - which variety or species shall increase in number, and which shall decrease, or finally become extinct.

The tree of life should perhaps be called the coral of life, base of branches dead; so that passages cannot be seen-this again offers contradiction to constant succession of germs in progress.

I cannot persuade myself that a beneficent and omnipotent God would have designedly created parasitic wasps with the express intention of their feeding within the living bodies of Caterpillars.

Man scans with scrupulous care the character and pedigree of his horses, cattle, and dogs before he matches them; but when he comes to his own marriage he rarely, or never, takes any such care.

Nothing before had ever made me thoroughly realise, though I had read various scientific books, that science consists in grouping facts so that general laws or conclusions may be drawn from them.

But when on shore, & wandering in the sublime forests, surrounded by views more gorgeous than even Claude ever imagined, I enjoy a delight which none but those who have experienced it can understand.

I was a young man with uninformed ideas. I threw out queries, suggestions, wondering all the time over everything; and to my astonishment the ideas took like wildfire. People made a religion of them.

When the views entertained in this volume on the origin of species, or when analogous views are generally admitted, we can dimly forsee that there will be a considerable revolution in natural history.

If Mozart, instead of playing the pianoforte at three years old with wonderfully little practice, had played a tune with no practice at all, he might truly have been said to have done so instinctively.

Blushing is the most peculiar and the most human of all expressions. Monkeys redden from passion, but it would require an overwhelming amount of evidence to make us believe that any animal could blush.

It has been a bitter mortification for me to digest the conclusion that the "race is for the strong" and that I shall probably do little more but be content to admire the strides others made in science.

I have steadily endeavoured to keep my mind free so as to give up any hypothesis, however much beloved (and I cannot resist forming one on every subject), as soon as facts are shown to be opposed to it.

It is absurd to talk of one animal being higher than another...we consider those, where the intellectual faculties most developed as the highest. - A bee doubtless would [use] ... instincts as a criteria.

May we not suspect that the vague but very real fears of children, which are quite independent of experience, are the inherited effects of real dangers and abject superstitions during ancient savage times?

I have steadily endeavored to keep my mind free so as to give up any hypothesis, however much beloved (and I cannot resist forming one on every subject), as soon as the facts are shown to be opposed to it.

Whoever is led to believe that species are mutable will do good service by conscientiously expressing his conviction; for only thus can the load of prejudice by which this subject is overwhelmed be removed.

Ignorance more frequently begets confidence than does knowledge: it is those who know little, and not those who know much, who so positively assert that this or that problem will never be solved by science.

Thus disbelief crept over me at a very slow rate, but at last was complete. The rate was so slow that I felt no distress, and have never since doubted even for a single second that my conclusion was correct.

If I had life to live over again, I would give my life to poetry, to music, to literature, and to art to make life richer and happier. In my youth I steeled myself against them and thought them so much waste.

When I view all beings not as special creations, but as the lineal descendants of some few beings which lived long before the first bed of the Cambrian system was deposited, they seem to me to become ennobled.

The loss of these tastes [for poetry and music] is a loss of happiness, and may possibly be injurious to the intellect, and more probably to the moral character, by enfeebling the emotional part of our nature.

I believe we were all glad to leave New Zealand. It is not a pleasant place. Amongst the natives there is absent that charming simplicity .... and the greater part of the English are the very refuse of society.

I have been speculating last night what makes a man a discoverer of undiscovered things. As far as I can conjecture the art consists in habitually searching for the causes and meaning of everything which occurs.

There are several other sources of enjoyment in a long voyage, which are of a more reasonable nature. The map of the world ceases to be a blank; it becomes a picture full of the most varied and animated figures.

People complain of the unequal distribution of wealth [but it is a far greater] injustice that any one man should have the power to write so many brilliant essays... There is no one who writes like [Thomas Huxley].

There is no fundamental difference between man and the higher mammals in their mental faculties...The difference in mind between man and the higher animals, great as it is, certainly is one of degree and not of kind.

We may confidently come to the conclusion, that the forces which slowly and by little starts uplift continents, and that those which at successive periods pour forth volcanic matter from open orifices, are identical.

If it could be demonstrated that any complex organ existed, which could not possibly have been formed by numerous, successive, slight modifications, my theory would absolutely break down. But I can find no such case.

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